Hamilton (1964) outlined two ways in which kin selection altruism could be favoured.
Firstly, if individuals have the capacity to recognize kin (kin recognition) and to adjust their behaviour on the basis of kinship (kin discrimination), then the average relatedness of the recipients of altruism could be high enough for this to be favoured. Because of the facultative nature of this mechanism, it is generally regarded that kin recognition and discrimination are unimportant except among 'higher' forms of life (although there is some evidence for this mechanism among protozoa). A special case of the kin recognition/discrimination mechanism is the hypothetical 'green beard', where a gene for social behaviour also causes a distinctive phenotype that can be recognised by other carriers of the gene. Hamilton's discussion of greenbeard altruism serves as an illustration that relatedness is a matter of genetic similarity and that this similarity is not necessarily caused by genealogical closeness (kinship).
Secondly, even indiscriminate altruism may be favoured in so-called viscous populations, i.e. those characterized by low rates or short ranges of dispersal. Here, social partners are typically genealogically-close kin, and so altruism may be able to flourish even in the absence of kin recognition and kin discrimination faculties. This suggests a rather general explanation for altruism. Directional selection will always favor those with higher rates of fecundity within a certain population. Social individuals can often ensure the survival their own kin by participating in, and following the rules of a group.
from the web page encyclopedia.thefreedictionary.com/Kin+selection
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