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Yes there is a difference. They are physically the same size casing, but the 556 military rounds' chamber pressure is ~10,000 psi higher then 223. Military rifles generally have chambers with a different throat configuration than normal sport 223 rifles. This being said, I would not recommend shooting 556 unless your rifle was designed for the higher pressures.

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Q: Is there a difference between 223 caliber and 556 mm?
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