The E symbol means 'times ten to the power of'.
So 4E7 means 4 x 107 which is 40,000,000.
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Calculator is feminine in French. The French word for calculator, calculatrice, ends in an e. Most words that end in e in French are feminine. Not always, but most of the time, you can rely on that rule.
It means Error. If you try to divide a number by 0 it should give you the E. Also if you try to multiply numbers that will give a larger number than the calculator allows. I believe exponential calculators fix this by giving you a exponential answer.
e-KT
20000000000000000
If you have a scientific calculator, there should be a key to input numbers in scientific notation. This key may be labeled "E" (uppercase E - the lowercase "e" means something different) or "Exp". In this case, press 1, EXP, 12 (meaning, 1 x 1012).