Oh, dude, the last letter but one is the second-to-last letter in the alphabet. So, like, if you're talking about the English alphabet, it would be "Y." But, you know, if you're talking about another alphabet, then it would be whatever comes before the last letter. Easy peasy, right?
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From my point of view, I think this is a trick question, so it can't be 'e', because that would be too easy to guess. Wait a second...you're asking a question, so it can't be a sentence! Is that correct? (I might be wrong :), so don't blame me for being slow in the brain.)
Last letter but one means, The second last letter or we could say like second letter from the last.
For Example consider a word "Dialogue". Here last letter but one is 'U'.